After witnessing a mech refused medical aid because of his class, Ratchet turned his back on everything he once knew, instead becoming the Elite that dared to change his place in an effort to fix what he knew was wrong. Little did he realize that walking into the small medbay would violate the very laws of society and threaten the dangerous, violent mechs he calls brothers.

Optimus Prime has been killed, Earth destroyed, and the universe on the verge of destruction, but the Allspark will do anything in its power to reverse time. Sam Witwicky is now the unwilling hero sent to Cybertron thousands of years in the past to redo a wrongdoing and to ensure that above all else, Optimus Prime must survive.