Joran Vydros, an Ex-Turian Legionnaire and former mercenary, becomes a candidate for the Spectres after being blackmailed by his family. At first he believes the Spectres to be nothing but corrupt; however, he later discovers being a Spectre to mean much more. After becoming Captain of the PFS Serpentine, he, alongside other comrades, discover a plot that will see a galaxy divided.

The conflict between the Empire and the Revolutionary Army rages on. A general from the southwest is returning to the capital, who could change the tides of war. While assigned to the Capital, the city awakens an ancient trauma that had been dormant in the recesses of his mind. He struggles with his past and the love for an old friend.